2307 AD. As fossil fuels became exhausted, humanity found a new energy source to change their situation: A large-scale solar power system with three enormous orbiting elevators. However, only a few large countries and their allies reaped the benefits. Three superpowers had ownership of the three orbiting elevators: The Union, based in the United States Of America, The People`s Reform League, made up of China, Russia, and India, and Europe`s AEU. The superpowers continue playing a large zero-sum game for their own dignity and respective prosperity. Even though it is the 24th century, humanity has yet to become one. In this world where war has no end, a private militia appears advocating the eradication of war through force. Each possessing a mobile suit Gundam, they are Celestial Being. The armed intervention by the Gundams against all acts of war begins.
Nena Trinity mourns the death of her brothers. Meanwhile on the moon Alejandro Coroner becomes incresingly agitated by Aeolia Schenberg's foresight. The crew of the Ptolemaios finish repairs to Dynames and pick up another GN Arms from its asteroid base within the Laragne 1 resource asteroid belt. The crew discuss the Trans-Am system, commenting how it provides a boost to performance, but only lasts a short time before the GN Drive must replenish the spent energy. Setsuna and Lasse make their way back to Ptolemaios and detect UN carrier ships heading to the Meisters' location. Warned of the attack, Allelujah and Tieria deploy in their Gundams, but Lockon is prevented from leaving. The battle soon commences between the Gundams and GN-X along with Ali Al Sarchez in the Gundam Throne Zwei. Allelujah and Tieria activate their Trans-Am systems and quickly destroy numerous GN-X. However, they become overwhelmed when their Trans-Am systems power down. Relief appears in the form of Dynames docked with its GN Arms system. Lockon uses the weapons platform to destroy more GN-X and two of the carrier ships before it is destroyed by Ali. Lockon chases after Ali determined to avenge his dead family. Zwei and Dynames engage in a vicious battle and Ali manages to cripple Lockon's Gundam. While Lockon makes Haro pilot the suit back to Ptolemaios, he exits the Gundam with his targeting rifle. Using an intact GN Cannon from his destroyed GN Arms, Lockon succeeds in destroying Zwei, but a shot from Zwei also damages the Cannon. As Lockon floats nearby he thinks about the world, its state and how he dilikes the current world. Holding his hand out to the world he questions those living on the Earth, if they like the current world because he doesn't. The GN Cannon explodes killing Lockon. Celestial Being suffer their first devestating loss leaving Setsuna to scream into the depths of space.
In the final confrontation between Setsuna and Ribbons, the 00 Raiser is heavily damaged by Ribbons and the other Innovators; however, assistance from the other Meisters even the odds. In the ensuing chaos, Allehujah and Hallelujah work together to defeat Hilling Care, while Lockon narrowly manages to kill Revive Revival. Eventually, Ribbons and Setsuna damage each other to the point where neither one can continue fighting. However, Ribbons manages to steal one of the 00's GN Drives, which was 0 Gundam's, and places it in the abandoned 0 Gundam. Setsuna, in turn, takes the remaining GN Drive and installs it back into the Exia. After another difficult duel, Ribbons and Setsuna run through each other with their swords but Setsuna emerges the victor. In the epilogue, the Federation undergoes reforms as its new president disbands A-Laws and pushes for better global peace; Saji and Louise rekindle their relationship; Allelujah and Marie have left Celestial Being to search for the meaning of their existence; Andrei Smirnov stays in the military to preserve peace like his father and mother; Patrick Colasour and Brigidier-General Kati Mannequin marry; Klaus and Shirin work as politicians in the Federation Assembly; Marina returns with the children to lead a rebuilt Azadistan; Graham Aker is seen visiting Billy Katagiri in his office; Lockon and Setsuna leave with Celestial Being (who are still considered dangerous due to their refusal to disband) to space and Tieria disappears into Veda to watch over the world.
Setsuna continues to struggle against Alejandro's weapons and shield. Exia docks with the GN Arms and evens the odds with the Alvatore. Setsuna succeeds in destroying Alejandro's shield but the GN Arms is destroyed in the process. Meanwhile, Sergei and Soma search for Kyrios. Allelujah wishes to know the meaning of their fight and unites with Hallelujah to face the two GN-Xs. Allelujah destroys Soma's GN-X and cripples Sergei but does not finish them off when he recognizes Soma as someone from his past. With his mobile armor damaged, Alejandro emerges in a golden mobile suit. Setsuna uses Exia's 'trump card' (his physical GN Blade) to bypass Alejandro's GN Shield and defeat him. Alejandro gets a message from Livonze revealing that Alejandro was a pawn and that Livonze is the true mastermind, right before Alvatore explodes. Graham then appears in his GN Drive-equipped Flag and engages Setsuna. Setsuna and Graham finally reveal themselves to each other and destroy each other's mobile suits. On Earth, Marina reads a letter from Setsuna questioning the current state of the world and how he hoped she'd help him learn the answers. Setsuna lies unconscious as Exia floats in space. Four years later (A.D. 2312), Saji writes a letter to Louise from space and says he is still waiting for her. He sees a GN particle trail in the distance and recognizes it as a Gundam. The UN has become the Earth Sphere Federation and all countries have disarmed and formed the ESF peacekeeping force. On Celestial Being's asteroid base, Ian talks with Wang Liu Mei about deploying their new mobile suits. Liu Mei asks to see the O Gundam, the first generation unit, and then comments that the OO Gundam will change the world.
Three new mobile suits, piloted by a trio of siblings, appear on the battlefield and reveal themselves as Gundam Throne Eins, Zwei, and Drei. Caught off-guard by the new Gundams' sudden arrival, the Union, AEU, and HRL forces are decimated and forced to retreat. In order to secure a route of escape, Gundam Throne Drei covers the entire region in GN particles, causing havoc with enemy communications and preventing a counterattack. After escaping from the Taklamakan Desert, the rescued Gundam Meisters are anxious about the new units and disturbed about not knowing of their existence. While world leaders despair at the appearance of the new Gundams, Alejandro Corner attends an anonymous conference with the other Celestial Being "observers" to review Celestial Being's activities and to discuss the acknowledgement of the Gundam Thrones. Aboard their spaceship, the Trinity siblings fly to meet the Ptolemaios to greet the Gundam Meisters.
It has been four months since Celestial Being's first intervention and the Gundam Meisters have been involved in over sixty intervention operations. As the world begins to change around them, the crew aboard the Ptolemy and the Gundam Meisters have some time to themselves.
Setsuna contemplates his memories of Lockon, who vouched for him when he first became a Meister. Tieria blames Setsuna for Lockon's death, but Sumeragi reminds him that their true enemies are still nearby. Meanwhile, having lost over half their GN-X and the Throne Zwei, Sergei suggests the UN force withdraw. However, UN headquarters reveals that reinforcements are coming and orders a second attack. Back on Ptolemaios, the Gundams undergo repairs and upgrades. Tieria suggesting they destroy all the pseudo-GN Drives so that they can resume their original plan. The rest of the crew contemplate their pasts, but an alarm signals that the UN forces have arrived. Alejandro Corner joins the UN forces in a large, golden Mobile Armor and fires a massive particle cannon at the Ptolemaios, which barely evades the blast. With Kyrios and Nadleeh protecting the ship, Setsuna and Lasse attack Alejandro but can't penetrate his GN Shield. Alejandro continues to fire on Ptolemaios causing massive damage. Tieria activates his Trans-Am system and destroys several GN-X, including Patrick Colasour's GN-X, but suffers extensive damage. A GN-X manages to approach Ptolemaios and fires directly into the bridge resulting in the death of Christina and Lichtendahl. Sumeragi, Ian and Feldt manage to escape in one of Ptolemios' containers. Lasse rams Exia's container into Alejandro's shield and breaks through. Exia and the GN Arms escape the destroyed container and charge back at Alejandro.
Setsuna and Saji head out to Lagrange 5 to locate Wang Liu Mei, who has, along with Hong Long, miraculously survived the attack by the Throne Drei. Nena intercepts the pair and shoots Hong Long, who sacrifices himself to allow Wang Liu Mei to escape. Liu Mei encounters Setsuna, and after giving him the coordinates to Veda, is killed by Nena as she tries to escape.
The duel between Setsuna and Graham ends with Setsuna emerging victorious. However, he chooses not to finish Graham off and leaves. Setsuna and the others soon review Wang Liu Mei's information and determine that a fierce battle against A-Laws will ensue as they journey to Veda's core at L2.
After dealing with Regene Regetta's treachery by having Ali-Al Saachez shoot him, Ribbons Almark reveals his massive colony ship, which he claims is the place where Aeolia Schenberg created Veda, the original GN Drives, and the Innovators. The colony fires a massive particle cannon and destroys most of A-Laws fighting combatants. Celestial Being launches an assault on the Innovator's colony ship and are countered by an army of Bring Stabity and Devine Nova clones, all piloting Trans-Am equipped kamikaze units. In the ensuing battle, Patrick Colasour sacrifices himself to defend Kati's ship and the Seravee is completely disabled by Healing Care and Revive Revival in their newly Trans-Am equipped suits. In the meantime, Setsuna and Saji confront Louise and Andrei; after disabling Andrei's Ahead, Setsuna moves in on the Regnant and Saji contacts Louise. The Ptolemaios II manages to enter the colony under cover from the Arios and GN Archer, but the ship is quickly infiltrated by hordes of automatons. Lockon fights a losing battle against Ali-Al Saachez in his rebuilt Arche Gundam, while Soma's GN Archer runs out of particles and is hit by an Innovator kamikaze. Sumeragi takes a gun and heads out to face the automatons swarming the Ptolemy, but is shocked to find they were accompanied by Billy Katagiri, the supplier of the Innovators' Trans-Am systems, who then pulls a gun on Sumeragi. Inside Veda's core, Ribbons is shocked that Tieria managed to survive Seravee's near-destruction and is now facing him, to which Tieria then pulls a gun on Ribbons.
As Lockon recovers from his injuries, the UN Army announces its plan to destroy the Gundams, "Operation Fallen Angel". The HRL's Choubu Squadron continues to harass the exhausted Trinity siblings. Observing on Ptolemaios, the Meisters wonder if they must be destroyed to finally end war. Setsuna rejects this notion, stating that the actions of both the Thrones and the U.N.
In the course of the Ptolemaios II encounter with an A-Laws task force, Revive is captured after his encounter with Tieria. Saji and Louise meet in battle, the two being transported to an alternate reality and Louise is left shaken in her resolve to fight against him. While Sumeragi and the Meisters interrogate Revive, his presence on the ship causes Anew to reveal her allegiance to the Innovators and declares the hijacking of the Ptolemaios II as she fires her gun at Lasse Aeon. Wang Liu Mei and Hong Long try to leave Ribbon's hideout to deliver Veda's location to Celestial Being, only for their ship to be shot down by Nena Trinity with Gundam Throne Drei.
To break his stalemate against Mr. Bushido, Setsuna takes an extreme risk by activating the 00 Gundam's Trans-am system in order to win. Soma's Ahead and the Arios are shot down and declared MIA as the Ptolemaios 2 is forced to retreat. Stranded and forced to face each other outside their mobile suits, Soma suffers mental attacks from Allelujah's presence and reverts to her Marie personality. Upon finding them, Sergei eventually decides to let them go and the two are rescued by Lockon. Meanwhile, Marina and Katharon learn the Federation intends to reorganize the Middle East and has dissolved all nations including Azadistan.
After the Memento Mori is fired at the Sairu Kingdom, Katharon begins planning an attack to destroy the superweapon. Tieria, after witnessing the devastation, decides to tell his comrades about the Innovators. When A-Laws forces led by Kati launch an attack on Celestial Being's asteroid base, everyone within is forced to flee. During the ensuing battle, Ian Vashti is injured, leaving Saji to deliver the 0-Raiser to the 00 Gundam. The combined unit, the 00 Raiser, is able to dispatch the A-Laws attacking, but when Setsuna activates the Trans-Am system, he and everyone in the area are affected by the particles and Saji and Louise shocked to learn they are fighting on opposing sides.Watch Now:Amazon
Anew reveals her allegiance to the Innovators and with Revive attempts to steal 0 Raiser for Ribbons but the machine is retaken, though the Innovators escape. Later they return, with new Suits (including Louise in her Regnant). Lyle attempts to get through to Anew but before he could, he is caught off guard, as Ribbons taken control of Anew. But as she was about to strike him down, her suit is shot by 00 Raiser as the Trans-Am particles have them share one tender moment together before her suit explodes.
As Setsuna attacks the three Gundam Thrones, he is surprisingly assisted by Tieria and Lockon. Tieria reveals Gundam Nadleeh's Trial System can disable any Mobile Suit linked with Veda. However, the Thrones manage to escape after informing Lockon that Setsuna was once a child soldier of the terrorist organization that was responsible for the death of Lockon's family. Lockon confronts Setsuna, but Setsuna's explanation of his past, as well as his conviction to end warfare as a Gundam Meister, defuses the situation. Meanwhile, Louise asks Saji to return to Japan to fulfill his dream as an aerospace engineer and to move on remembering the memories as dreams, and Kinue discovers that "Ragna" refers to the president of the corporation constructing linear elevators for the Orbital Elevators. Meanwhile, as Alejandro and Livonze continue their mysterious agenda, the three superpowers, about to surrender before the Gundams, are contacted by a member of Celestial Being to reveal a facility in Antarctica housing 30 GN Drives.
Saji mourns the loss of his sister. The Trinity siblings are betrayed and launch a brief counterattack after their compound's location is revealed, destroying one of the 10 GN-X units from Choubu's assault force. The UN Army continues its assault on Celestial Being, attacking the Ptolemaios in space. Raguna Harvey is killed after Corner learns that it was he that created and sold the GN-X units. The Meisters' units are out-numbered, and they struggle to defend themselves against the GN-Xs firepower. During the battle, Alejandro Corner accesses Veda and deactivates the GN drive of the Meisters' units, leaving them defenseless. However, Sumaragi activates a backup system and every unit except Virtue is able to reactivate. Seeing Tieria's crippled state, Patrick Colasaur goes for the killing blow. Lockon is able to intercept the attack, but is severely injured. The battle ends with the UN Army retreating after losing two of their GN-X units, and the timely arrival of the GN Arms.
Marina speaks to Setsuna, Tieria, Ian and Lasse, thanking them for saving her while making a request for them to be bring her back to Azadistan when Tieria refused at first. On the way to Azadistan, the Ptolemaios 2 is attacked by an A-LAWs navy ship in an ambush and has deployed underwater mobile armor 'Trilobyte' to eliminate the ship. Setsuna, Lockon, Tieria and Allelujah have managed to defeat them, bringing the fight to the surface before Soma and The Masked Man intercepted Setsuna and Allelujah with their customized Aheads. Kataron mobile suits arrive just in time to drive back the A-LAWs ship. Marina and Shirin are then reunited. Meanwhile, Billy has been recruited as an A-LAWs MS development chief due to his grudge with Sumeragi.
With assistance and supplies from Katharon, the Ptolemaios 2 is able to continue with its repairs in the mountainous region. However, they are unable to proceed with their plan to break through the siege laid by the Federation, without the currently missing Setsuna and the 00 Gundam. Meanwhile, Setsuna, wounded after his fight with Ali Al-Saachez, is forced to get medical help at Katharon's Suille base, where he is reunited and cared for by Marina. The Ptolemaios 2 is forced to defend itself against an overwhelming number of A-Laws mobile suits, but receives an unexpected respite when the fleet suddenly retreats from its successful offensive to deal with a coup d'état led by Hank Hercule at the African orbital elevator. Despite his injuries, Setsuna sets off towards the elevator and is met by Mr. Bushido, who forces Setsuna to fight against him by demonstrating the Masurao's secret ability: Trans-Am.
The Gundam Thrones continue to attack military installations around the globe, their aggressive tactics forcing the Celestial Being observers and the Gundam Meisters aside and out of the forefront of battle. The crew of the Ptolemaios investigate the Thrones and their origins, including the possibility that Veda was hacked by an outsider to acquire the technology behind the GN Drive. On the civilian forefront, Kinue Crossroad obtains a new lead after learning that one of the Throne pilots mentioned something called 'Ragna'.
Surviving Katharon guerrillas, including Shirin and Marina, go to negotiate with the Sairu Kingdom, the only country in the Middle East with considerable military power to oppose the A-LAWs. Meanwhile, Sumeragi leads Celestial Being in returning back to their asteroid base, where Setsuna retrieves the 0-Riser. A subsequent test run of the 00 Gundam's twin drive system reveals the potential of the unit. When the A-LAWs' new orbital weapon fires a particle beam on the Sairu Kingdom's capital, Allelujah's Hallelujah personality returns and Marie senses that Sergei is in trouble.Watch Now:Amazon
A-Laws attempts to retake the Africa Tower using assault automatons to kill both the rogue troops and civilian hostages alike, although the Innovators continue to manipulate the information released to the public in order to conceal the A-Laws' brutality. Meanwhile, Setsuna is locked into a heated battle with Mr. Bushido and his Masurao. Their battle is interrupted with the arrival of the Ptolemaios 2 and the remaining Gundam Meisters. After analyzing the unusual redeployment of Federation troops around the elevator, Sumeragi and Kati both realize, to their horror, that they are anticipating that the elevator will collapse soon and that A-Laws is planning on completely destroying the Africa Elevator. A renewed Memento Mori superweapon is then shown preparing to fire at the orbital space elevator.
After the Meisters land on the damaged Kataron base in the Middle East, they were greeted with accusations and distrust. At this time, Tieria brings Saji back to the Ptolemaios 2 after he spotted him all shaken up in guilt. A-LAWs launches another operation to sortie against the Ptolemaios 2. The Meisters launch to intercept while Sumeragi is out of action after a flashback of a failed operation she led that resulted in her love interest, Emilio's, death. Saji follows Ian to a Ptolemaios 2 turret, while initially hesitating to do so before he fires it at a GN-X III piloted by Louise Harlevy. Allelujah encounters difficulties while facing Soma and Setsuna with Mr. Bushido. As a result, Setsuna goes into trans-am mode with his 00 Gundam.
During his encounter with Regene Regetta, Tieria learns the Innovators' views on VEDA and Aeolia Schenberg while the A-LAWs obtain two new pilots: Patrick Colasour and Innovator Revive Revival. Tieria and Setsuna go undercover to infiltrate a party with A-LAWs and Earth Federation officials, where Tieria meets with Ribbons Almark, who reveals that according to Aeolia's plan, Tieria should have died with the rest of Celestial Being four years ago. Refusing an offer to join the Innovators, Tieria and Setsuna, who has been exposed to Louise as a member of Celestial Being by Billy, escape in their Gundams.
Setsuna escorts Marina back to Azadistan after the Meisters stop at Kataron's Middle Eastern base. Saji leaves the base as he does not trust the Celestial Being's action. Suddenly, an Earth Federation ship arrives in the area and detains Saji before Sergei decides to release him. An A-LAWs task force arrives with Mr. Bushido, Soma and Barrack leading the squad before automaton droids were deployed to massacre Kataron guerrillas and civilians. Lockon arrives with Tieria and Alelujah to hold back the A-LAWs task force with Lockon gunning down the droids in rage while Soma hesitates to attack him when she sees the droids attack the guerrillas and civilians. Setsuna and Marina arrive in Azadistan, only to face Ali with a mobile suit as he watches the country burning.
After entrusting Lyle Dylandy with top secret information pertaining to Celestial Being's activities, Setsuna goes to confront Sumeragi Lee Noriega who, consummed by guilt over the countless lives lost during the organization's military interventions four years prior, found refuge in Bill Katagiri and worsened her drinking condition. Meanwhile, Wan Liu Mei leaks information to Ribbons and Saji Crossroad is given access to Celestial Being's archives and learns about the existence of the Throne Gundams. A-LAWS forces mount an attack on Celestial Being's ship, the location of which was among the data acquired by Ribbons, and Tiera is left alone to defend the Ptolemaios. Setsuna, accompanied by Lyle and Sumeragi, heads towards the ship and boards the 00 Gundam, easily dispatching the attacking units and saving the Ptolemaios.